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Staff+ Application Security Engineer

Anthropic· Security
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Hybrid (Remote-friendly with travel; SF, Seattle, NY offices)$320k - $485kfull-timePosted 9mo ago

About the role

Senior security engineering role focused on building Claude-powered application security systems, threat modeling, and secure design reviews for Anthropic’s AI-powered product stack.

Responsibilities

  • Design, build, and operate Claude-powered security systems — LLM-driven code analysis, automated vulnerability remediation, AI-assisted threat modeling — and own one or more of them end-to-end, including the cross-functional relationships that come with it
  • Lead secure design reviews and threat modeling for novel AI systems, identifying risks that don't map to existing frameworks
  • Evolve a public bug bounty program where automation handles routine triage and root-cause work, and engineers handle escalations and corner cases
  • Partner with Product, Infrastructure, and Research teams as an embedded security owner — consulting on launches, shaping architecture, and influencing decisions where security is the constraint
  • Share an operational on-run rotation with the rest of the team — bounty escalations, incident response, and launch consults on systems serving Claude in production

Qualifications

  • Hands-on application and infrastructure security experience, including cloud and containerized environments
  • Production-quality coding ability in at least one of Python, Go, Rust, or TypeScript, with a track record of building durable systems rather than one-off scripts
  • Practical threat-modeling and vulnerability-identification skills — you've found and reasoned about real bugs in real systems
  • Demonstrated ability to operate with high autonomy and ambiguity
  • Clear technical communication with both engineers and leadership

Skills

Required:Hands-on application and infrastructure securityProduction-quality codingThreat modeling and vulnerability identificationAutonomy and dealing with ambiguityTechnical communication

Benefits

  • Competitive compensation
  • Optional equity donation matching
  • Generous vacation and parental leave
  • Flexible working hours
  • Office space for collaboration
Last updated Jul 8, 2026. Interested? Apply on the company site.

About Anthropic

building reliable, interpretable, and steerable AI systems

Anthropic is an AI safety and research company that's working to build reliable, interpretable, and steerable AI systems.

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