Principal Engineer, AI Security
Cambridge, MA$252k - $374kfull-timePosted 2w ago
About the role
Senior individual contributor responsible for security engineering of Lila Sciences' AI/ML systems, driving AI security controls across data, models, pipelines, and agent runtimes, and ensuring continuous compliance and risk reduction throughout the AI lifecycle.
Responsibilities
- Turn the AI security roadmap into delivered controls with measurable outcomes
- Design and implement security controls across training and inference pipelines, model registries, MCP servers, and agent runtimes
- Keep AI infrastructure continuously compliant against internal standards and applicable frameworks
- Drive vulnerability remediation across AI infrastructure, models, and dependencies to closure with system owners
- Reduce risk across data, models, pipelines, and agentic systems through threat modeling, control design, and adversarial assessment
- Partner with AI Platform, AI Research, ML Operations, Science, and AI Safety so security is embedded at design time
Qualifications
- Strong background in security engineering, cloud security, or DevSecOps with hands-on experience securing production infrastructure
- Working knowledge of AI/ML systems and the ML lifecycle, plus AI-specific risks such as prompt injection, model and data poisoning, insecure agent tooling, and inference-time threats
- Experience with software supply chain security and dependency and vulnerability management
- Experience implementing data security controls (classification, data flow mapping, access control, encryption) in cloud environments
- Proven ability to partner across engineering teams and drive cross-functional remediation to closure
- Comfort operating with ambiguity in a fast-moving research environment
Skills
Required:Security engineeringAI/ML lifecycle knowledgeSoftware supply chain securityData security controlsCross-functional collaboration
Benefits
- Full-time U.S. employees receive medical, dental, and vision coverage; employer-paid life and disability insurance; flexible time off with generous company-wide holidays; paid parental leave; educational assistance program; commuter benefits including bike share; company subsidized lunch program
- International Benefits tailored to region; USD salary ranges apply to U.S.-based positions; international salaries set to local market
Last updated Oct 7, 2026. Interested? Apply on the company site.
About Lila Sciences
Pioneering Scientific Superintelligence
Pioneering Scientific Superintelligence to solve humankind’s greatest challenges.
Scientific SuperintelligenceAIAutonomous LabsLife SciencesChemical SciencesMaterials Sciences
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